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Authors: Imlay, Ralph W.
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Date: /1959
Title: Succession and Speciation of the Pelecypod Aucella
314-G , , 155-167 , pls. 16-19
Host: U.S. Geological Survey Professional Paper:
Document Type: Article\Abst.\Chapter\report
Annotation: ABSTRACT

The pelecypod Aucella, of latest Jurassic and earliest Cretaceous age, has been subdivided into more than a hundred species on the basis of mirnor differences without allowing for biological variation, or for variation due to crowding that is normal for an attached gregarious pelecypod. As a result, in any large collection from a single locality, specimens can be selected that fit the definitions of a number of species and yet these so-called species are connected by many specimens showing transitional characteristics. Such an assocition appears, therefore, to represent a single variable population. If the species of Aucella are defined making allowance for variation, the number of species is greatly reduced, but the specific concepts can be grasped readily by geologists and used for mapping purposes. Seven zones based on species of Aucella can now be recognized easily in the field in North America, as has been demonstrated by geologists of the U.S. Geological Survey.
UserNote
L.M. No. 3 [L.M. is Lawrence Martin] (Mesozoic loc. 3124) , Mt Katmai , West side of Katmai Bay 2 1/2 mi. SW. of village. [Note by RBB: Martin, 1926, USGS Bull. 776, p. 211 gives following description: "West side of Katmai Bay 2 1/2 miles southwest of village, or half a mile south of lagoon. Talus of dark sandy shale that is somewhat higher than the beds that yielded lot 3101. Lawrence Martin, 1904."]; [West side of Katmai Bay 2 1/2 miles southwest of village, or half a mile south of lagoon. Talus of dark sandy shale that is somewhat higher than the beds that yielded lot 3101. Lawrence Martin, 1904. (description from Martin, 1926, p. 211)]; [USGS Mes. Cat.: 3124. Orig. no. 3. Jurassic. W. side of Katmai Bay, 1/2 mile N. of camp, Shelikof Region, Alaska. Coll. L. Martin, Aug. 13, 1904. N.B. 2, p. 29.] [Note discrepancy in loc. description with that given in Martin, 1926]; Lawrence Martin, 1904. West side of Katmai Bay, 0.8 km north of camp on west side of Shelikof Strait, Mt. Katmai Quadrangle, on Alaska Peninsula. Naknek Formation. Late Oxfordian to early Tithonian. (description from Imlay, 1981, USGS PP 1190, p. 27); [Entry in USGS Mesozoic Catalogue (Washington, D.C.): 3124. Orig. no. 3. Jurassic. W. side Katmai Bay, 1/2 mile N. of camp, Shelikof Region, Alaska. Coll. L. Martin, Aug. 13, 1904, N.B. 2, p. 29.]
Aucella spitiensis Holdhaus
L.M. No. 5 [L.M. is Lawrence Martin] (Mesozoic loc. 3127) , Mt Katmai , West shore of Katmai Bay 1/2 mi. N. of 3124 [Note by RBB: Martin, 1926, USGS Bull. 776, p. 212 gives following description: "West side of Katmai Bay. Talus half a mile north of 3124 and at about the same horizon. Lawrence Martin, 1904."]; [West side of Katmai Bay. Talus half a mile north of 3124 and at about the same horizon. Lawrence Martin, 1904. (description from Martin, 1926, p. 212)]; [USGS Mes. Cat.: 3127. Orig. No. 5. Jurassic. S. shore of Katmai Bay, 1 mile N. of camp, Shelikof Straits Region, Alaska. Coll. L. Martin, Aug. 13, 1904. N.B. 2, p. 29.]
Aucella spitiensis Holdhaus
E of C.N. Fenner, 1919 (Mesozoic loc. 10248) , Mt Katmai , Naknek formation, about 500 feet above sea level on Bear Mountain, Katmai Bay, Alaskan Peninsula, Alaska. C.N. Fenner, 1919. (description from Imlay, 1959, p. 166); [Highly fossiliferous stratum on Baer Mountain, near Lagoon Camp, Katmai Bay, north of camp and about 500 feet above sea level. C.N. Fenner, 1919. (description from Martin, 1926, p. 212)]
Aucella concentrica (J. de C. Sowerby)
No Data (Mesozoic loc. 11360) , Unknown , Naknek formation, Fossil Creek, north of Lake Becharof, southwest Alaska. Dr. Laymore, 1922. (description from Imlay, 1959, p. 166)
Aucella rugosa (Fischer)
47AGr239 (Mesozoic loc. 21028) , Mt Michelson , Kingak shale, Canning River, 2 1/2 miles south of Black Island, lat 69 deg. 30' 45" N., long 146 deg. 18' 45" W., northern Alaska. George Gryc, 1947. (description from Imlay, 1959, p. 166); Many USGS Mesozoic localities near the Canning River, northeast Alaska. Mesozoic locs. 24014, 21028, 25598, and 29882 (Imlay, 1955, p. 79, 80; Keller and others, 1961, p. 193) from lat 69°30'45" N. to 69°33' N.; long 146°18' W. to 146°23' W.; Mesozoic locs. 29134-29136 near Shublik Island at lat 69°24' N., long 146° 10' W. Amoeboceras (Prionodoceras) and Buchia concentrica (Sowerby). (description from Imlay and Detterman, 1973, p. 25); [USGS Mes. Cat.: 21028 (orig. #47AGr239). Jurassic - Kingak. N.E. 1/4 N.E. 1/4 682, lat. 69°30'45", long. 145°18'45", 2 1/2 miles south of Black Island, Canning River area, Alaska. Black shale with limestone interbeds immediately underlying Leffingwell's Ignek (Upper Cretaceous) formation, (about same position as collection Gr.8f and 32F). Gryc - August, 1947]; George Gryc, 1947. West bank of Canning River. 4 km south of Black Island, lat 69°30'45" N., long 146°18'45" W., SE. cor. sec. 19, T. 2 N., R. 24 E., Mt. Michelson Quadrangle. Black shale with limestone interbeds near top of Kingak Shale. Late Oxfordian to early Kimmeridgian. (description from Imlay, 1981, USGS PP 1190, p. 23); George Gryc, 1947. Canning River, 2 1/2 miles south of Black Island, NE1/4 NE1/4 quad. 682, lat 69°30'45" N., long 146°18'45" W. Black shale containing ironstone interbeds near top of Kingak shale. Upper Jurassic, upper Oxfordian or lower Kimmeridgian. (description from Imlay, 1955, USGS PP 274-D, p. 80)
Aucella spitiensis Holdhaus
49ASa43 (Mesozoic loc. 22127) , De Long Mts , Tiglukpuk formation, large cutbank on southwest side of Kukpowruk River, lat 68 deg. 42'30" N., long 163 deg. 14' W., northern Alaska. E.G. Sable, 1949. (description from Imlay, 1959, p. 166); USGS Mesozoic loc. 22127. Kukpowruk River, lat 68°42'30" N., long 163°14' W. (Imlay, 1955, p. 78,79). Buchia rugosa (Fischer). (description from Imlay and Detterman, 1973, p. 26); [USGS Mesozoic Catalogue (Washington D.C.) entry: 22127. Orig. no. 49ASa43F. Lower Cretaceous - Killik Fm., northern Alaska, Wainwright district, NW 1/4 SE 1/4 Quad. 686. Kukpowruk River. Sandstone, shale, concretions. Coll. Edward G. Sable. N.B. Sable p. 32, June 1949]; E. G. Sable, 1949. Kukpowruk River, approximate lat 68°42'30" N., long 163°14' W., eastern part of a large cut bank on southwest side of river. Sandstone and shale containing concretions. Stratigraphic position unknown. Upper Jurassic, Kimmeridgian to lower Portlandian. (description from Imlay, 1955, USGS PP 274-D, p. 78)
Aucella rugosa (Fischer)
50ALz6 (Mesozoic loc. 22472) , Misheguk Mtn , M.C. Lachenbruch and A.H. Lachenbruch, 1950. North bank of Seagull Creek, Nimiuktuk Valley. DeLong Mountains. Lat 68 deg. 19'50" N., long 159 deg. 59' W. Okpikruak formation, lower part. Berriasian. (description from Imlay, 1961, USGS Prof. Pap. 335, p. 21)
Aucella okensis Pavlow
50AKt113 (Mesozoic loc. 22514) , Howard Pass , B.H. Kent, 1950. Northeast of Ekakevik Mountain. Lat 68 deg. 36' N., long 156 deg. 57' W. Gray to green graywacke. Okpikruak formation. Valanginian. (description from Imlay, 1961, USGS Prof. Pap. 335, p. 24)
Aucella crassicollis Keyserling
51AMo24 (Mesozoic loc. 22724) , Misheguk Mtn , R.H. Morris, 1951. Ridge one quarter of a mile west of lakes at the base of Thunder Mountain. Lat 68 deg. 40' N., long 160 deg. 20' W. Sandstone lenses in shale. Okpikruak formation, middle part. Berriasian. (description from Imlay, 1961, USGS Prof. Pap. 335, p. 20)
Aucella okensis Pavlow
51AMo156 (Mesozoic loc. 22736) , Misheguk Mtn , R.H. Morris, 1951. East bank of Nuka River. Lat 68 deg. 41' N., long 160 deg 08' W. Limestone and sandstone lenses in shale. Okpikruak formation, lower part. Valanginian. (description from Imlay, 1961, USGS Prof. Pap. 335, p. 20) [description given in Imlay, 1959, gives slightly differing lat. (68 deg. 40'30" N) and long. (160 deg. 03' W)]
Aucella sublaevis Keyserling
51AKe23 (Mesozoic loc. 22757) , Sagavanirktok , A.S. Keller, 1951. Kemik Creek. Lat 69 deg. 26' N., long 147 deg. 10' W. Siltstone lenses. Okpikruak formation, lower 700 feet. Berriasian. (description from Imlay, 1961, USGS Prof. Pap. 335, p. 29)
Aucella okensis Pavlow
51AKe77 (Mesozoic loc. 22762) , Chandalar , A.S. Keller, 1951. Kashivi Creek. Lat 69 deg. 12' N., long 147 deg. 44' W. Siltstone. Okpikruak formation. Berriasian. (description from Imlay, 1961, USGS Prof. Pap. 335, p. 29)
Aucella subokensis Pavlow
51AKe153 (Mesozoic loc. 22768) , Philip Smith Mts , Kingak shale, from 500 to 800 feet above local base at Lupine River, lat 68 deg. 52' N., long 148 deg. 22' W., northern Alaska. Samuel Keller, 1951. (description from Imlay, 1959, p. 166); USGS Mesozoic Iocs. 22750, 22751, 22766, 22768, 22769. Lupine River from lat 68°49'30" N. to 68°51' N., long 148°17'30" W. to 148°22'30" W. (Keller and others, 1961, p. 195). Buchia rugosa (Fischer), B. mosquensis (von Buch), and B. concentrica (Sowerby). (description from Imlay and Detterman, 1973, p. 26); [USGS Mes. Cat.: 22768. Alaska -51AKe153 - Kingak - 5'-800' base - sta. no. K-155 - 682- NE of SW quad. - Lupine River - 68°52' N - 148°18' W - siltstone]
Aucella rugosa (Fischer)
54AKe154 (Mesozoic loc. 22769) , Philip Smith Mts , Kingak shale, about 300 feet above local base at Lupine River, lat 68 deg. 52' N., long 148 deg. 22'W. (description from Imlay, 1959, p. 166); USGS Mesozoic locs. 22750, 22751, 22766, 22768, 22769. Lupine River from lat 68°49'30" N. to 68°51' N., long 148°17'30" W. to 148°22'30" W. (Keller and others, 1961, p. 195). Buchia rugosa (Fischer), B. mosquensis (von Buch), and B. concentrica (Sowerby). (description from Imlay and Detterman, 1973, p. 26); [USGS Mes. Cat.: 22769. Alaska - 51AKe154 - Kingak - 300' above Triassic contact - sta. no. K -157-682. NE of SW quad. - Lupine River - 68°52'N - 148°17' W - siltstone and shale]; A. S. Keller, 1951. Lupine River, lat 68°52' N., long 148°22' W. Shale as at Mes. loc. 22766. About 300 feet above base of Kingak shale. Upper Jurassic, middle Kimmeridgian. (description from Imlay, 1955, USGS PP 274-D, p. 80)
Aucella mosquensis (von Buch)
51ASa75 (Mesozoic loc. 22780) , Misheguk Mtn , E.G. Sable, 1951. Hills three-fourths of a mile east of east-central fork of Driftwood Creek. Lat 68 deg. 37'15" N., long 160 deg. 31'25" W. Okpikruak formation, probably lower part. Valanginian. (description from Imlay, 1961, USGS Prof. Pap. 335, p. 19)[Imlay, 1959, p. 166 gives slightly differing lat/long data: 68 deg. 38'05" N., long 160 deg. 29'20" W.]
Aucella crassicollis Keyserling
51ASa255 (Mesozoic loc. 22790) , Misheguk Mtn , E.G. Sable, 1951. North bank of the Nuka River. Lat 68 deg. 42' N., long 160 deg 00'45" W. Interbedded sandstone, siltstone, and silty claystone. Okpikruak formation, 25 feet above base. Berriasian. (description from Imlay, 1961, USGS Prof. Pap. 335, p. 20)[Imlay, 1959, p. 166 gives slightly different lat/long - 68 deg. 42'10" N., long 159 deg. 56'50" W.]
Aucella okensis Pavlow
51ASa287 (Mesozoic loc. 23554) , Misheguk Mtn , E.G. Sable, 1951. North bank of Nuka River. Lat 68 deg. 40'30" N., long 160 deg. 05'20" W. Shale containing limestone lenses. Okpikruak formation, about 300 feet above base. Valanginian. (description from Imlay, 1961, USGS Prof. Pap. 335, p. 21)[Imlay, 1959, p. 166 gives slightly different lat/long - 68 deg. 40'45" N., long 160 deg. 02'45" W.]
Aucella sublaevis Keyserling
51ASa1045 (Mesozoic loc. 23560) , Misheguk Mtn , E.G. Sable, 1951. North bank of Nuka River. Lat 68 deg. 41'30" N., long 160 deg. 02'15" W. Sandstone and shale containing limestone lenses. Okpikruak formation. Valanginian. (description from Imlay, 1961, USGS Prof. Pap. 335, p. 20)[Imlay, 1959, p. 166 gives different longitude - 159 deg. 59' W.]
Aucella crassicollis Keyserling